MadSci Network: Environment |
First, I am a real person, although it is sometimes safer to write using a pseudonym, so having satisfied that I'll turn to the food web. Even the temperate grassland biome doesn't exist as a single type of community. A number of different types of grasslands exist, and a complete food web is probably not known for any of them. Although I cannot answer in the specific, I can describe the general characteristics of a food web in such grasslands. The term food refers to eating and being eaten, biological facts of life. Autotrophs, literally self-feeders, can capture energy from environmental sources and use it to synthesize complex molecules from simple raw materials. The energy that organisms use is stored in the chemical bonds of these molecules. Not all organisms can do this, so all the rest, heterotrophs (other feeders) must obtain complex raw materials and energy from some other organism, either an autotroph or another heterotroph. However, ultimately all the energy available to organisms is the sum total captured by the autotrophs. Many different species compose both categories, and the complex patterns of eating and being eaten form a food web. In temperate grasslands, the predominate autotrophs are photosynthetic, largely flowering/fruiting plants (Angiosperms)in the Grass family. Members of the Bean, Sunflower, and Mint families also make considerable contributions with many species among the common plants. So food webs start with green plants. They use the energy they captured for growing and reproducing. Heterotrophs consist of two large categories, consumers who eat their food, and decomposers who largely grow upon or in their food obtaining food by absorption. Herbivores are primary consumers, those who eat the autotrophs. In grasslands a number of mammals function as herbivores, and they usually fall into two categories, small, no bigger than a rabbit or prairie dog, and big, like deer, antelope, and bison. The reason for this discontinuous distribution is that plants like grasses are of low nutritional value with a lot of bulk, and it takes a big animal to graze upon such food. A lot has to be comsumed and a long, slow GI tract is needed to digest it. Smaller mammals can select among an array of plants, selecting those with the highest nutritional value, but since these are a smaller and patchier resource, smaller mammals are better adapted for using this resource. Even deer are pretty select eaters on grasslands and their browsing is far from random. Although the mammals may be the most obvious to another large mammal like ourselves, the largest group of herbivores in number, diversity, and total mass will be insects. Some will be specialists on particular plants, others will be more generalists. In some instances, seed-eating birds and some reptiles can be common herbivores too. The remaining heterotrophs are seconday consumers, basically carnivorous predators who consume the primary consumers, the herbivores. Birds, ranging from small insect eaters to large mammal eaters (hawks, falcons) are common grassland predators. Much rarer are larger predators, usually members of the Dog (coyote, wolf, fox), Cat, and Weasel families. Large predators are rare for two reasons. First, as energy moves from one trophic level to the next, from plants to herbivores, not all the energy captured is available because some was used for the plant to grow and reproduce, and the transfer of energy isn't completely efficient. As a general rule, no more than 10% moves to the next trophic level, so the total biomass of herbivores is never more than 10% of the total biomass of plants. Predators would be less than 10% of the consumers, and the reduction in habitat size further prohibits their populations. In terms of energy, it takes a pretty big area of grassland to support bison, and even more to support large predators like wolves. So as a general rule, big, fierce animals are always rare in comparison to herbivores. Of course this isn't to suggest there is a straight interaction between grass, bison, and wolves. What makes these feeding interactions a web is that they are interconnected. Several different insects may feed on one species of plant, but not on another, which is largely eaten by a specialist. Insects also prey upon other insects, and then you add in the insect eaters, and the organisms that may prey upon them, and the resulting diagram with lines connecting organisms that eat and get eaten in turn begins to resemble a web or net. Lastly of course, all the waste products and bodies become energy sources for the decomposers. It would take years of study to document all such interactions in any particular grassland community. And even after years of working in such communities, we find new interactions all the time.
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