MadSci Network: Botany
Query:

Re: Why is xylem bigger then phloem?

Date: Thu Nov 4 18:42:38 2004
Posted By: David Hershey, Faculty, Botany, NA
Area of science: Botany
ID: 1099574287.Bt
Message:

Why questions are often difficult to answer especially in this case given that 
xylem and phloem function in very different ways. Thus, diameter of one is not 
necessarily related to the diameter of the other.

One possible reason is that xylem vessels are dead at maturity so are somewhat 
like tiny pipes. The Poiseuille law, also called Hagen-Poiseuille law, for 
laminar flow in pipes states that the rate of flow is proportional to the 
fourth power of the pipe radius. That means that one large diameter pipe 
provides much more flow than several smaller diameter pipes with the same 
total cross sectional area as the larger pipe. 

Xylem vessels have thicker walls than phloem sieve tubes because they must be 
able to withstand substantial tensions. The xylem sap is usually under tension 
during the day according to the prevailing cohesion-tension theory. Phloem is 
not subject to such great stresses. 

The size of mineral nutrient ions would not have anything to do with it. They 
are all very small compared to sugars, phytohormones, and other organic 
molecules that move in phloem.


References


Hydraulic properties of individual xylem vessels of Fraxinus americana


Calculating Velocities of Water Movement in the Xylem and in Living Cells

Rice, Stanley A. and McArthur, John. 2004. Water Flow Through Xylem: An 
Investigation of a Fluid Dynamics Principle Applied to Plants. American 
Biology Teacher Volume: 66: 120-127



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