MadSci Network: Botany
Query:

Re: Why photosynthesis pigment of plant is green and not black?

Date: Fri Dec 21 20:47:32 2001
Posted By: David Hershey, Faculty, Botany, NA
Area of science: Botany
ID: 1008863837.Bt
Message:

The first website cited describes an interesting idea that plants are green 
because purple Halobacterium evolved before chlorophyll-containing organisms. 
The photosynthetic Halobacterium absorbed green light so other organisms 
possibly evolved chlorophyll to absorb nongreen wavelengths and fill an 
ecological niche. The first two websites both argue that a black chlorophyll 
might absorb too much radiation and either overheat the plant or harm the plant 
by absorbing destructive UV and x-rays. 

It is important to remember that leaves often absorb more than half the green 
wavelengths and use them in photosynthesis. It is a widespread misconception 
that leaves reflect all green light. That misconception is based on looking at 
a chlorophyll absorption spectrum (see second website cited), which is obtained 
by extracting chlorophyll into an organic solvent, such as acetone, and 
measuring its absorption in a spectrophotometer at wavelengths between 400 and 
700 nanometers. The chlorophyll absorption spectrum does show that chlorophyll 
in a test tube absorbs only about 2 to 3% of the green light. However, that is 
very artificial because a leaf is highly structured. Salisbury and Ross (1985) 
note that in the intact leaf, a green photon may not initially be absorbed by a 
particular chlorophyll molecule but it is reflected and then gets another 
chance to be absorbed, and perhaps another, and another, etc. within the 
complex leaf structure that does not exist in a test tube. Thus, each green 
photon has many opportunities to be absorbed in the leaf so the total 
absorption of green light by chlorophyll is much higher in the leaf than in a 
test tube of extracted chlorophyll. Accessory pigments, such as carotenoids, 
also absorb green light (see second website cited) and funnel the energy to 
chlorophyll.

Instead of a chlorophyll absorption spectrum, people should be looking at a 
photosynthesis action spectrum (see third and fourth websites cited) which 
shows the amount of photosynthesis at each wavelength. Because plants do absorb 
substantial green and yellow wavelengths and use them in photosynthesis, plants 
are more efficient than they seem by assuming they only use the red and blue 
wavelengths that chlorophyll absorbs in a chlorophyll absorption spectrum. 
Photosynthesis is the most widely taught plant biology concept but it is often 
taught with numerous misconceptions.
 

References


Why Trees are Green


Photosynthesis Pigments


Photosynthesis Action Spectrum for Elodea


Generalized Photosynthesis Action Spectrum

Salisbury and Ross. 1985. Plant Physiology. Belmont, CA: Wadsworth.


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