MadSci Network: Botany |
Xerophytes have a variety of adaptations to reduce transpiration including fewer stomata, sunken stomata, thick cuticle, hairs that shade the leaf (as in old man cactus that looks like it is covered by gray hair), leaves that have evolved into spines with photosynthesis in the stem (as in cacti), CAM (crassulacean acid metabolism) in which stomates only open at night to fix carbon dioxide, and thick, fleshy leaves with greatly reduced surface area and water storage. Some plants reduce water loss in dry conditions by shedding their leaves. Ephemerals survive dry conditions as seeds and germinate, flower and set seed during a few weeks when the soil is moist. Window plants of the Kalahari desert have fleshy leaves with transparent areas on top. The leaves can be buried in sand and light can still reach down into the buried leaf. Having stomates on upper and lower leaf surface or just the lower surface does not seem to be an important distinction between mesophytes and xerophytes. References College botany text such as Stern, K.R. 1991. Introductory Plant Biology. Dubuque, Iowa: Wm. C. Brown Cronquist, A. 1971. Introductory Botany. New York: Harper and Row.
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